Fiona, struggling to adjust to life with her distant Aunt Tanis and her slimy boyfriend Dex after her parents' death, finds solace in an abandoned mansion filled with stray cats and birds. She discovered the mansion while skipping on summer dance classes. When the mansion is threatened with demolition, Fiona is determined to save it, seeing it as a symbol of finding a safe home, a feeling she deeply understands. However, the mansion's feline residents, led by the self-proclaimed "King" Cot, and prowling black cat Piper, have their own agenda, suspecting the newly arrived birds are behind the demolition threat.
As the demolition begins, Fiona seeks help throughout her community while the cats clash over their preferred methods of dealing with the situation. Cot wants to expel the birds, while Piper believes she can intimidate the demolition team. The birds, meanwhile, simply long to return to their original home. Fiona realizes that saving "The City" will require more than just her efforts; she must unite the disparate animals and rally her community to stop the luxury condo development and create a better future for everyone.
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